Ichthyosaurs

Ichthyosaurs painted by Heinrich Harder (1916)

History of Life
(Geology 100)
Department of Geology, The College of Wooster
Spring 2009

Preparation Questions

Latest Quiz Key

Sample Test

Geology Course Pages

Welcome to the webpage for the History of Life course at The College of Wooster! You will find here our syllabus (a "living syllabus" which is added to every week) and the details of the course itself. For each week's topics I post links, images and other items to provide additional information (and alternative views) as we explore the wonders of life's history through the semester. This page is essentially a required blog for the course, as well as a set of resources we will use in class. To keep our material as up-to-date as possible, I post supplementary information only a week in advance. I thus expect registered students in the course to visit this site at least once a week. If you have ideas about items you would like to see posted here, please send me e-mail (mwilson@wooster.edu) or talk to me in class. I hope you enjoy the course! You may also want to visit our "Invertebrate Paleontology at Wooster" general page, our Invertebrate Paleontology course page, and our Paleontology Independent Study webpages.

Mark A. Wilson
Department of Geology
The College of Wooster
Wooster, OH 44691 USA
mwilson@wooster.edu


Schedule of Lectures
(With Weekly Links to Other Web Resources)
 
Lectures are in Scovel Hall Room 105 on Mondays, Wednesdays and Fridays from 8:00 - 8:50 a.m.

January 12-16

Antiquity of the Earth and Life (Geologic Time and the Fossil Record)
Recommended Reading: Allaby pages 44-47; Prothero pages x-58
Special Events:
Assignments: (Preparation questions)

Web Resources for Week #1:

Geologic time is a fascinating topic. It is so unimaginably long it has been called "deep time". Please learn the Geologic Time Scale as soon as possible because it is the chronological language of this course. Here is a clever test you can take on the Web when you think you've learned the terms. If you want to see ALL the time terms geologists use, check out this PDF of the geological time scale. You need to know only the eras, periods, and Tertiary and Quaternary epochs. William Smith was an Englishman responsible in many ways for the principles behind our use of geologic time. James Hutton was a Scot who first effectively described and applied the concept of "uniformitarianism" in geology, which requires vast amounts of time. NOTE: Because the required terms of the Geological Time Scale for this class are unclear from the above links, I've posted a MicroSoft Word version of our time scale as well as our own pdf version. You will see that I recognize the Ediacaran Period before the Cambrian.

Geologists distinguish between relative and absolute time, with the first being the ordering of events, and the second using measurements of time. We will most often use relative time (such as the "Cambrian Period") in this course, but also occasionally cite absolute time ("65 million years ago").

The concepts of radioactive decay and its use in the dating of Earth materials take some time to master. The University of Waikato in New Zealand has an excellent explanatory website on radioactive dating. On this site called "popcorn" you will see an Applet display of decay demonstrating its spontaneity and constant rate. (It loads slowly but is worth it.) A more detailed explanation of how half-lives are really calculated is shown here. See what I have spared you?

As an introduction to the fun of paleontology, there are plenty of dinosaur animation sites on the Web. Try searching for "dinosaur" on YouTube and you'll get thousands of odd videos! (Including, of course, "Walk the Dinosaur" of happy memory.) You will also want to visit The Paleontology Portal which is "a central entryway to paleontology on the Web". It is an excellent link to thousands of paleontological resources. Of course, as a preview of what's coming, you simply must see the evolution of Homer Simpson!

You may not be surprised to learn that the scientific framework of this course is considered a lie or delusion by almost half of Americans. "Young Earth Creationists" believe that the Earth and the Universe are a few thousand years old, and that evolution did not occur. This is radically different from the cosmological and evolutionary models supported by scientists. Creationist arguments appear scientific, but you will quickly see that they are based on misconceptions, misrepresentations, mysteries, and zealotry. For example, take a look at this page citing "evidence for a young Earth" from Answers in Genesis. These are the people who produced the multimillion-dollar Creation Museum in Kentucky. Don't ever confuse their evangelism with science. The first part of the Prothero readings this week is an excellent analysis and history of the creationism movement in the United States. We will not be covering creationist arguments directly in this course (we have science to do), but I will always answer any questions you have about them. I am an evolutionary paleontologist, but please be assured that you will not be judged or graded on your personal beliefs. What you believe is always your business. What you understand about evolution and the history of life is mine.

Geology in the News:

February 12, 2009, will be the 200th anniversary of the birth of Charles Darwin (and Abraham Lincoln!). As part of the commemoration, we will see many summaries of the best evidence for organic evolution. Here is a very nice compendium (in pdf form) titled 15 Evolutionary Gems from the journal Nature. We will refer to this several times in class, so you might want to print a copy.

China is still a frontier for paleontology. Now it appears that the Chinese may have discovered the largest dinosaur bone deposit yet known. It is Late Cretaceous in age and contains at least 7600 bones in a single excavation pit. One of the fossils may be the largest hadrosaur (20 meters long!).

About 13,000 years ago there was a global cooling event which ushered in the Younger Dryas and is correlated with a loss of large mammals and the Clovis human culture, especially in North America. The causes have been fiercely debated recently. The latest evidence is an abundance of "nano-diamonds" in spherules thought to be derived from a meteorite impact. Note that some scientists in the article say such materials are not necessarily from an impact. [Note also this January 29 follow-up article which casts more doubt on this impact hypothesis.]

A team drilling a hydrothermal well in Hawaii actually punched down into a magma chamber. This is a geological dream because while we see lava flows often, we rarely get a chance to sample magma still below the surface. One geologist said, "It's the difference between looking at dinosaur bones in a museum and seeing a real, living dinosaur roaming out in the field."


January 19-23

Mechanisms of Organic Change: Evolution
Recommended Reading: Allaby pages 52-53; Prothero pages 86-119
Special Events:
Assignments: (Preparation questions)

Web Resources for Week #2:

This week we begin discussing evolution. (The "e word" as a high school administrator once told me as he was trying desperately not to use it.) Start with a good definition of evolution, for you will soon see that dorm room conversations on the topic often have significant errors and misconceptions. You can then visit "Frequently Asked Questions About Evolution" at TalkOrigins Archive. The University of California Museum of Paleontology at Berkeley has an excellent website on the history of evolutionary theory, including good sections on pre-Darwinian thinkers. The Berkeley pages on "Understanding Evolution" are the best on the web. Charles Darwin (again, February 12, 2009, is the 200th anniversary of his birth) is well represented web-wise, and Alfred Russel Wallace is the subject of this excellent site by Charles Smith of Western Kentucky University. The full text of Darwin's most famous work, The Origin of Species (which appeared 150 years ago), is available on-line. It is not scintillating reading, at least not on a computer, but you may want to peruse the preface and the conclusions.

The public opposition to evolutionary theory is deep and strong in this country, and it has spread internationally. Anti-evolutionists are very diverse, and many of their beliefs are contradictory. Here is a graphical and textual classification of beliefs on origins and evolution. "Scientific Creationism" and other anti-evolution movements are founded on nonsense, ignorance, wishful thinking and a deep undercurrent of anti-intellectualism, but they are part of democracy's fabric and the public discourse on science. We should listen to them closely and try to understand the social and political forces behind their strong beliefs (and yet still vigorously teach evolution, of course). Some creationist websites are sophisticated, such as that of the Institute for Creation Research and Answers In Genesis (AiG). (Several years ago I debated Mr. Ken Ham of AiG on a radio talk show in New York City. That was an experience.) These sites and others provide access to the primary tenets of scientific creationism. I especially recommend the questions-and-answers page of AiG and the Handy-Dandy Evolution Refuter. The National Center for Science Education (NCSE) is the best known defender of evolution in American public schools. The Creation-Evolution Reference Database is a good place to start for anyone who wishes to study this topic in detail, although it is not updated.

For an insight into the political implications of anti-evolutionism, you can't do better than to read the opinion of Judge John E. Jones in the recent Dover, Pennsylvania, "Intelligent Design" court case (a long pdf).

This excellent computer graphics animation of DNA replication and protein synthesis is spectacular. Thank you, Mícheál O'Duffy, for finding it. It hits all the major points we discuss in lecture.

Geology in the News:

"Clouds of Methane May Mean Life on Mars." Or not. Methane is most often produced on Earth by life processes, but there are inorganic reactions which can also generate methane. I think this is a way over-hyped story from those seeking more funding for "astrobiology". Still, we've learned something new about Mars: some process on or in it is making seasonal clouds of methane.

Here's yet another dinosaur found with preserved feathers. And it is once again in China, which has been a source of some amazing fossils in the last decade. In this case the dinosaur is Beipiaosaurus, a small theropod. The feathers were clearly for insulation and maybe display, but not for flight. Few people doubt any more that birds are evolutionarily derived from dinosaurs -- or may even BE dinosaurs!

From recent discoveries in Israel, it appears that our ancestor Homo erectus could not only control fire in some way, but at least 790,000 years ago could make fire with flints. This pushes back the known date for producing fire at least 500,000 years.

Human predation in the form of hunting and fishing is causing rapid evolution in prey species. This is the first time such an effect on tasty wild animals has been demonstrated. There is fear that this "dangerously fast" evolution will "disrupt ecosystems". No doubt it will, as evolution always does.


January 26-30

The Very Beginning: Origin of the Universe, the Early Earth & Life
Recommended Reading: Allaby pages 16-43; Prothero pages 144-159
Special Events:
Assignments: (Preparation questions)

Web Resources for Week #3:

The Big Bang Hypothesis is an extraordinary concept. Readable (and brief) explanations of Big Bang ideas can be found at "Stephen Hawking's Universe", and another much more extensive set of resources at "Hot Big Bang". You will want to especially see the page on the awesome singularity. Want an artist's view of the Big Bang? For you humanities majors, how about Big Bang Philosophy? And it wouldn't be the web without a site like "Why the Big Bang is Wrong!" For those of you looking for a simple red shift explanation, click away on this PBS webpage. It even has exercises you can try. There is a very cool Big Bang animation on this French website; I may use it in class (despite the French). I'm not sure what the point is of this next Big Bang animation, unless it is to make us feel insignificant.

This star formation site is good, but also very technical. The JPL-NASA Solar System site has the latest news and fantastic images. Some exquisite photographs of the Moon's surface are also posted by NASA. This webpage from NOVA has superb animations of the "Big Whack" hypothesis for the origin of the Moon. This Moon-formation video on YouTube is also excellent, although the music could be improved!

Neil Comins, an astronomer in Maine, wrote an interesting little book called "What if the Moon Didn't Exist?" The Moon is critical to the maintenance of life on Earth, which I find amazing. Also, here is the homepage of NASA's "Stardust" project, which brought to Earth the "dust" of a comet. Now you know the importance of this study to our ideas about the origin of the solar system because comets are samples of the original solar nebula.

Now for the complex and contentious ideas about the origin of life. The elaborate and elegant Exploring Life's Origins website from the Museum of Science in Boston is the best on the topic. Our friends in the chemistry department at Beloit College have this origins webpage with animations of the Miller-Urey experiment. (Here's the nice animated Miller-Urey experiment webpage I may use in class.) The Creationist opposition to all naturalistic theories for the origin of life is characteristically strong, but their slick webpages have numerous errors. Don't forget those panspermians who avoid the issue of origins altogether -- they say life came from space, "where it always was"!

If you are interested in being on the e-mailing list for the Department of Geology, please send me a note. Our list has the names of students who enjoy geology, maybe to the point of considering it as a major or minor. You will receive departmental announcements about upcoming lectures, picnics, field trips, and internship opportunities

Geology in the News:

Your fellow student Mícheál O'Duffy sent in this recent story about the fossil "hobbits" found on the Indonesian island of Flores several years ago. The debate has been whether these organisms are members of our own species or separate hominin primates. This skull analysis concludes they are another species. Very interesting. I bet we'll have another view by the end of the semester!

A paper just published in the Zoological Journal of the Linnean Society demonstrates that Cretaceous birds had brains just as advanced as those of today's birds. This observation is advanced as a possible explanation for why birds survived the Cretaceous extinctions and their dinosaur cousins did not. I'm not so sure, especially since the boundary between birds and dinosaurs has been almost completely erased.

I just found this volcanism blog which tracks volcanic eruptions around the world. It is always impressive to see the concurrent volcanic activity around the world, most of which we hear little about.


February 2-6

The First Record: The Oldest Fossils
Recommended Reading: Allaby pages 48-51; Prothero pages 160-171
Special Events:
Assignments: (Preparation questions)

Web Resources for Week #4:

Fourth week already? We will begin with continued discussions of origins, especially hypotheses about the origin of life. Please be sure to look at the links listed for last week on this topic. We will also examine the earliest fossil record, which goes back at least 3.5 billion years. The University of Arizona has a nice page on the six kingdoms of life, which makes a good starting point. Another website to read would be the one on fossil bacteria at the University of California, Berkeley. This site also has an excellent page on cyanobacteria and stromatolites, which are the first easily-recognized fossils. Check out their Archean page, though, and find the critical error! (It is in the first paragraph.) Here is another stromatolite page. Queen's University in Ontario also has a good early life online museum website. There is chemical evidence for life which even predates the oldest stromatolites. For an excellent short summary of "the emergence of complex life", you can't beat this colorful page from the University of Michigan.

Here is a recent news article I want to bring up into our discussion: "In Our Genes, Old Fossils Take On New Roles", from the September 1, 2008, edition of the Washington Post. Note how the "central dogma" of inheritance is stretched by the work retroviruses can do on our genetic composition. "It turns out that about 8 percent of the human genome is made up of viruses that once attacked our ancestors." There are enormous implications here for how evolution works. This is one of the reasons why the standard evolutionary tree is beginning to look a bit strange with its new intersecting branches.

We are now beginning to order geological and evolutionary events in time. You might want to refer to this annotated timeline as a reference. Do not take it too literally, though, because there are some events either not recorded or now known to have occurred at different times. Again, the events you should be most concerned about are those we cover directly in class

Geology in the News:

Mícheál O'Duffy found a YouTube animation of DNA replication and protein synthesis. The graphics are extraordinary, and it is just about at the level we discussed in class. It is so good that I will add it to our course material. Just remember as you watch this: it is computer graphics!

Just to show what you find when you examine a problem with a hypothesis, scientists are discovering that the genetics and physiology of the obscure placozoans shows that they are similar to the earliest animals. It is not a surprise to learn "that evolution does not proceed along a straight line", but it is very useful to flesh out, as it were, the creatures at the base of our family tree.

Analysis of fractures in their preserved skulls suggests that the dinosaur Triceratops "likely engaged in horn-to-horn battles with its kin." Very much like rams, which is not surprising given the trend in the last two decades to find that dinosaurs behaved more like birds and mammals than today's reptiles.

Remember that idea of a mammoth-killing impact event postulated to have happened 12,900 years ago? A recent analysis of charcoal and pollen in the pertinent sediments shows that it likely didn't happen. I was skeptical from the first!

More evidence that Charles Darwin was on the side of the angels: his fierce anti-slavery beliefs may have played a critical role in his development of the modern theory of evolution. This is from a new book by Adrian Desmond and James Moore titled "Darwin's Sacred Cause".

Huge volcanic eruption likely soon from Alaska's Mount Redoubt. As the historian Will Durant cogently wrote, "Civilization exists by geological consent, subject to change without notice."


February 9-13

Developing Marine Ecosystems
Recommended Reading: Allaby pages 58-59; Prothero pages 172-197
Special Events:
Assignments: First Lecture Exam (February 13) (Preparation questions)

Web Resources for Week #5:

Our topic this week is the rise of animal life on Earth, and the early development of marine communities. A good place to start on the Web is Berkeley's "Vendian Animals" page with its many illustrations and links (in class we will refer to this time interval as the "Ediacaran"). Queen's University of Canada also has a good Ediacaran fauna page. (Canadians are big on the earliest rocks for good reason -- they have a lot of them!) There is also a very professional Ediacaran Page from an amateur in New Zealand. His Cambrian and Ordovician pages are excellent as well.

We will spend considerable time on the rise of skeleton-bearing invertebrates in the Cambrian. Please visit Berkeley's Cambrian page, especially their subpage on the "Tommotian" (the earliest Cambrian). I hope you will be fascinated with the Burgess Shale (this is the Smithsonian's main page on it), found in the Middle Cambrian. This is the Smithsonian's Burgess Shale reconstructed images page. I am very fortunate to be able to visit the Burgess Shale quarries in British Columbia this coming summer. A site called "Prehistoric World" has some great little Burgess Shale animations. The Cambrian also was the most important time for those fascinating little critters, trilobites. You can join the Yahoo Trilobite Club if you like and have access to an extraordinary number of images and links for these extinct animals. The "Palace of Trilobites" (these people are VERY enthusiastic) is in Japanese, but the universal binomial nomenclature and beautiful images will help you out. There are many more of these out there in cyberspace.

For some photographs and brief studies of Ordovician fossils in the Cincinnati area, you can't do better than Wooster's own "Keck Ohio" pages (although they are getting a bit dated in this fast-paced webworld). You can also see Wooster paleontology students collecting Ordovician fossils in Indiana last semester. The Palaeos webpages on the Ordovician are very well done and updated often.

Here is sample test #1 for your preparation. Remember that the test will begin at 7:55 a.m. on Friday. Don't forget to know the Geologic Time Scale so well you can recite it in your sleep. It will be the last problem on your test -- guaranteed.

Geology in the News:

We have a new record for the oldest animal fossils: chemical evidence of demosponges in Oman approximately 635 million years old. Chemical fossils, in this case a set of steroids called steranes, are a bit tricky because they can be implanted by contamination, but these results look very robust.

A huge quake in China last year may have been triggered by a massive dam project. The earthquake in Sichuan was a magnitude 7.9, and it killed thousands and left millions homeless. Could the massive Zipingpu dam and reservoir have caused the temblor? It is a difficult question because the quake itself is natural but the timing could have been accelerated by the weight of water and concrete.

Maiacetus, the "good mother whale", is an excellent transitional fossil in the lineage of terrestrial mammals leading to aquatic cetaceans. These animals still had their hips in place. What is cool about a recent fossil find of Maiacetus is that it has a fetus preserved within its skeleton.

A fossil snake has been found which is estimated to have been approximately 13 meters long and weighing over 1100 kg (2500 pounds). Titanoboa may have eaten crocodiles for breakfast! I don't think I mind not having the chance to meet this monster!


February 16-20

Plate Tectonics
Recommended Reading: Allaby pages 82-105, 140-149
Special Events:
Assignments: (Preparation questions)

Web Resources for Week #6:

Our primary topic this week is the Theory of Plate Tectonics, with an emphasis on how the dynamic Earth has affected the evolution of life. We can start with this beautiful world map, showing continents, mountains, ridges, transform faults and trenches in the Pacific and Atlantic Oceans. Here are some maps of the ancient Earth, with helpful color coding. This site (from an astronomical observatory in South Africa) has a good long page summary of plate tectonic theory. (It is a bit slow, but worth the wait.) NASA can show us by satellite laser ranging data the modern rates of plate movement. (One of my brothers used to work on this project.) The Nevada Seismological Laboratory at the University of Nevada, Reno, has an excellent page on plate tectonics and earthquakes. This is an extraordinary site, complete with on-line research articles. The University of Texas hosts The Plates Project which concentrates on producing accurate plate reconstructions of the past. Of course, plate tectonics is an ideal subject for web animations. PBS reviews plate tectonic theory with a simple and clear set of images and animations. Here is a global history of continental movement in the last 750 million years from UC Berkeley. The University of Kentucky has some very cool animations of the last 150 million years you control with the cursor. This page also takes a few seconds to load because of the sophisticated graphics.

As for the primary effect of plate tectonics on us, visit the real-time global earthquake website maintained by the U.S. Geological Survey. You will see how very common earthquakes are around the world, and where you do not want to buy a house. PBS has a nice set of animations of earthquakes from a global perspective, along with volcanoes and the geological phenomenon we now know all too well, tsunamis. (Here is a summary page on the devastating 2004 tsunami in the Indian Ocean.)

This week we also begin a discussion of the causes of the Permian Mass Extinctions -- the greatest disaster in the History of Life. Paleogeographic maps of Pangaea at the end of the Permian can be found on the Scotese website (note that it was not "99% of all life" which went extinct) and on the Permian Page from Northern Arizona University. Wikipedia has a good Permian Extinctions article, although the topic is controversial enough that some parts of it may be edited by partisans of one cause or another.

Here is the electronic version of your essay assignment which will count as two quizzes. Remember that it is due on Friday, February 27th.

Geology in the News:

On Darwin's 200th birthday, Gallup Poll released the results of a survey of Americans on how many "believe in evolution". By now you are not shocked to see that only 39% of Americans say that they do. I think that word "believe" is a mistake and that there are other ways to ask about acceptance of evolutionary theory, but the fact that most Americans are either in opposition to or ignorant of evolution is a deep national shame.

When Fox News ran the above story, they also linked to another of their reports on "missing links and the fossil record", showing how common it is to find fossils which are "transitional" between groups. I think this was a very progressive thing to do on their part, and maybe just a few more evolution-deniers will think again.

CNN has a very good story on Darwin's Birthday, looking back at the man and the century and a half since the Origin of Species publication. There are good distinctions to be made between the evolutionary ideas developed by Darwin and the modern framework of the theory. There is a reason why scientists do not blithely refer to evolutionary theory today as "Darwinism". As with all good science, innovators first promote foundation-shifting ideas which are then modified and adapted to new evidence over time. A Darwinian process, I suppose!

Tell me what you think of this "experimental" video: the evolution of life in 60 seconds. "Blue-green algae" = cyanobacteria here. I'm not sure what "outgassing of molecules" means.


February 23-27

From Fins to Feet: Early Vertebrate Evolution
Recommended Reading: Allaby pages 56-57; Prothero pages 198-231
Special Events:
Assignments: Essay due on Friday, February 27, at 8:00 a.m. (Preparation questions)

Web Resources for Week #7:

Vertebrates! At last, our cousins appear (after we finish the extraordinary Permian Extinctions). The best place on the Web for simple and thorough descriptions of the chordates and their history is at the University of California Museum of Paleontology. Check out their pages on the urochordates (tunicates), cephalochordates, and chordates in general. The Berkeley tetrapod (four-legged) webpage is especially useful for us. This page on limb evolution from an anti-creationist website is very useful, even if the illustrations are a bit clunky. I like this website lovingly devoted to the ancient coelacanth fish Latimeria. It has many illustrations and an extensive history of what we know about this remarkable evolutionary survivor. The Tree of Life page on terrestrial vertebrates will show you current ideas about the relationships of fish and early amphibians. It is a rich source of further links. Our class lecture outline on the early vertebrates is found here. I'm giving this to you directly not to replace your notes but to give you a thorough and concise source for the concepts and vocabulary.

Here is a good general page from Norway on amphibians. Berkeley also has an amphibian page, but it is considerably shorter. If you're looking for a lot of frog images, look no further than this link. If you want to freak out your roommate right now, try amplifying a few frog calls from your computer.

Here is a link to the essay that is due this week on Friday, just in case you lost your paper copy.

Geology in the News:

If you're going to dig an underground parking lot next to the La Brea Tar Pits in Los Angeles, you're going to come across a lot of new Pleistocene fossils. The Page Museum at the tar pits is about to double its collection with this new material, which includes a nearly-complete mammoth and at least one sabretooth cat.

There are ideas afloat that life may have evolved more than once on Earth, and that some organisms may be around that are utterly unrelated to the rest of life as we know it. With our new genetic analysis tools we can begin the search for these so-called "alien" life forms among us (or even in us!).

Mount Redoubt in Alaska is still rumbling and will certainly erupt in the near future -- at least in geological terms. When is the question. You can keep track of it on the Alaska Volcano Observatory webpage.

If our skies clear up, we may be able to see Comet Lulin. On Monday it will be just south-southwest of Saturn in the southern skies. Well worth a look, especially with binoculars or a telescope.


March 2-6

The Amniotic Egg: Amphibians to Reptiles
Recommended Reading: Allaby pages 60-63; Prothero pages 232-247
Special Events:
Assignments: (Preparation questions)

Web Resources for Week #8:

The eighth week already. It must be time to really "conquer the land" as we say too often. The reptiles are an enormously successful class, and their diversity is reflected in the websites devoted to them. I can only give you links to a few of the many groups we will discuss in class this week. (Just wait until we get to the dinosaurs!) You will want to review the wonders of the amniotic egg. For the technically inclined, this is a complex page designed for those studying the amniotes. Here's an introductory page to the turtles (anapsid reptiles), and then another for the diapsids. One of our favorite synapsids is the reptile Dimetrodon. Grab this Dimetrodon image, a coloring page to print out (for your little brothers and sisters). Everything you want to know about the therapsids ("mammal-like reptiles") can be found here. To round out our initial survey of early reptiles, meet the "living fossil" tuatara. The best general reptile page I've found is from Heidelberg, Germany. It has many very informative (and richly illustrated) pages in English. I'm sure you'll like this mosasaur page from the South Dakota School of Mines and Technology Museum of Geology. I enjoyed this page on Cretaceous marine life from the University of Nebraska State Museum.

A pivotal event in Mesozoic reptile evolution was the Triassic mass extinction, which is well covered on this webpage from the University of Bristol. The rise of the dinosaurs starts soon after.

You need some silly links since this is the week before Spring Break. This is fruitcake pseudoscience, but you'd be surprised how many people believe these things. Some crackpots, for example, look at the evidence for plate tectonics and instead turn it into proof of an expanding earth? Crazy talk. These guys always sound desperate, craving scientific approval. An expanding earth, by the way, is the only way some people can explain the "surprisingly massive sizes" of dinosaurs. Here are some very nicely done videos of the expanding earth concept by a Neal Adams. You're supposed to give these to your geology professors to show what fools they've been. This crank is fond of satirical, angry letters to scientists, especially geologists, because they just don't seem to recognize his genius. Plate tectonics can't work anyway because the reptiloids of the inner earth wouldn't have their cozy underground caverns for millions of years. The Cretaceous extinctions, as a digression, were not caused by a natural disaster but were instead the result of a war between "two enemy alien groups". Here's something new -- a man working very hard to convince you with NASA photos that there are forests on Mars. All this time the government has been keeping this amazing news from us (while apparently still releasing the photos, silly Feds).

Again, our class lecture outline on the early vertebrates is found here. I'm giving this to you directly not to replace your notes but to give you a thorough and concise source for the concepts and vocabulary

Geology in the News:

Stephanie Jarvis sent in this link to a YouTube presentation on the origin of life. It is more detailed than what we could cover in class, but you will still enjoy it. Remember our prediction: we will have the basic outline of a unified origin of life hypothesis within a few years. This video shows how close we are.

Some gorgeous 1.5 million-year-old footprints of Homo erectus have been found in northern Kenya. They tell us much about the walking patterns and foot bone structure of the immediate ancestor to Homo sapiens. I'm impressed at how well the little toes show up!

Here is that story we talked about in class: a Devonian placoderm fish found with a tiny juvenile placoderm inside, which is apparently the earliest pregnancy in the fossil record. Note that this is not, as so many reporters write, the "earliest sex". That happened a long time before the Devonian, as you well know.

You've got to see this newly-found "psychedelic" fish found in Indonesia. So much more to be discovered out there!


March 23-27

Jurassic Park, etc.: The Evolution of Reptiles
Recommended Reading: Prothero pages 248-257
Special Events:
Assignments: (Preparation questions)

Web Resources for Week #9:

In this first week back from a glorious Spring Break (I'm sure), we'll start with therapsids and the Triassic mass extinction, which we didn't quite get to in week #8. (By the way, I learned that what I called in class the temporal fossae are more properly called the temporal fenestrae. Fenestrae means windows, and it refers to the openings. The fossae are depressions derived from the openings.) Then it is on to the big guys --

As you may imagine, the Web is rich with dinosaur pages. Some of these websites are excellent, some are horrible, most are tolerable (and long). Let's begin our explorations at the University of California Museum of Paleontology at Berkeley, which has by far the best dinosaur site. In fact, they invented the concept of the "on-line museum". Start with their general dinosaur page, which has numerous images and links. Then take a guided tour of the Berkeley Dilophosaurus exhibit and their "T. Rex Expo". I suggest ending your Berkeley dinosaur visit with their "Dinobuzz" page which has excellent explanations and discussions of the latest dinosaur research. Other impressive dinosaur websites include that of Paul Sereno, one of the most productive dinosaur paleontologists in the world and virtually an industry to himself. The "Journal of Dinosaur Paleontology" doesn't seem to be a real journal, but it contains some interesting discussions of dinosaur biology. Of course, I hope you've all seen the extraordinary series "Walking With Dinosaurs" on the Discovery Channel. The webpage for this series is superb.

The Dinosaur Interplanetary Gazette will keep you busy if you like everything dinosaurian. You might also want to "download-a-dinosaur" to make folded-paper dinosaurs for all your friends. (Notice, though, that some of the dinosaurs listed are not dinosaurs.) The National Geographic website has an excellent presentation on prehistoric life, including dinosaurs. We cannot forget, of course, all the wonderful evidence that dinosaurs and people lived together. (I hope my sarcasm is apparent.) How about that idea again that dinosaur size can be explained with an Expanding Earth? I love this craziness!

Here's another creationist page on dinosaurs you must see, beginning with this quotation: "This article will discuss the possibility that there may have been an ongoing effort since the earliest dinosaur 'discoveries' to plant, mix and match bones of various animals, such as crocodiles, alligators, iguanas, giraffes, elephants, cattle, kangaroos, ostriches, emus, dolphins, whales, rhinoceroses, etc. to construct and create a new man-made concept prehistoric animal called the dinosaur." So, apparently this David P. Wozney thinks scientists just made up dinosaurs.

Geology in the News:

Here's a complicated story sent in by Mícheál O'Duffy about a gene in humans which was "resurrected" from a non-functional past in earlier primates. There is a lot in this little story. I want you to simply note the complexities of a genetic inheritance, including the processes of genes being turned on and off and some genes "killing off" others.

National Geographic has a nice page showing some of the more charismatic "missing links" found since Darwin's time. Of course there have now been thousands of such "links" discovered throughout the fossil record.

Mícheál also gave me this link to high-resolution photoscans of the Iceman. These can be creepy, especially if you take a real close look into his shriveled eyes. The 3-D images are fantastic if you have those funky little red-blue glasses to wear. Ötzi, as he has been called, is about 5300 years old and was found frozen in a glacier on the border between Italy and Austria.

Creationists, including the "Intelligent Design" variety, were not invited to a Vatican conference on evolution. "We think that it's not a scientific prospective, nor a theological or philosophical one," said the Rev. Marc Leclerc, the conference director and a professor of philosophy of nature at the Gregorian. "This makes a dialogue very difficult, maybe impossible."


March 30 - April 3

Origin of Flight
Recommended Reading: Allaby pages 64-65; Prothero pages 257-269
Special Events:
Assignments: Second Lecture Exam (April 3) (Preparation questions)

Web Resources for Week #10:

Time flies, and now so do vertebrates. We will discuss the origins of vertebrate flight this week. The University of California Museum of Paleontology has an excellent set of pages on flight origins, which includes information on birds, bats and pterosaurs. For details on the latest pterosaur research, try The Pterosaur Database. For more popular questions, you can't beat the Pterosaur FAQs webpage. Wikipedia does a nice job with pterosaurs as well, although you always have to be a bit careful here. Stanford University and the National Geographic Society actually want to make a flying replica of a pterosaur. Here's a strange link: "evidence" that pterosaurs lived until the 17th Century, believe it or not. (I suggest "not").

The evolution of birds from reptiles is a fascinating topic, and there have been additions and changes to the story every year since this course began. You can start with this technical and well-illustrated account of bird evolution from the University of Guelph. Note, though, that many of the "unique avian features" are shared with dinosaurs. In the words of the Wikipedia page on bird evolution (which is very good), "As more non-avian theropods that are closely related to birds are discovered, the formerly clear distinction between non-birds and birds becomes less so." We will spend some time in class considering models for the origin of bird flight. You might want to visit UC Berkeley's high school page on bird flight evolution as well.

Here is a link to the second sample test. Your test will be different, I need not remind you.

Geology in the News:

Here's a good story sent to us by Rob McConnell concerning a "teenage gang" of Triceratops dinosaurs found preserved together. This is first-class evidence of dinosaur sociality which augments considerably what we now know about Triceratops.

Mícheál found for us this 360° view of a coral reef in New Caledonia. It takes a little while to load, but it is worth the vicarious journey across a modern reef with all its incredible diversity.

The Cambrian Burgess shale is back in the news with the discovery of a new fossil predator called Hurdia victoria. (I wish the headline writers didn't erroneously call it a "bug"!) The fossil specimens themselves were actually found a century ago, but clever paleontologists from Sweden and Canada pieced it properly together this year. Did I tell you I'm going to the Burgess Shale locality in August? Now everyone will want to go! Just like my upcoming visit to Svalbard and the new Predator X found there. I thought I was ahead of the crowd.

Mysterious domes on the surface of Mars may be mud volcanoes. This is not only evidence for subsurface water, but may also be a source for the methane gas detected in the Martian atmosphere. I'm still not enthusiastic about the possibilities for life on Mars.


April 6-10

Our Furry Kin (The First Mammals) and The Greening of the Earth: Evolution of Plants
Recommended Reading: Allaby pages 66-67; Prothero pages 270-282
Special Events:
Assignments: (Preparation questions)

Web Resources for Week #11:

Mammals evolved so gradually from the reptiles that it becomes difficult to say when the first mammal appeared. The Wikipedia page on the evolution of mammals is quite good, and the related page on the cynodonts (a type of therapsid which includes the mammal ancestors) is useful. We will talk in detail about the various innovations which characterize the mammal-like reptiles and the early mammals.

Plants have been in this course for awhile as a sort of leafy background to tetrapod evolution, but now it is time to take them seriously. The best place to start a general survey of our very distant green cousins is with Berkeley's fossil plants page. Their fossil plants range chart may be especially useful, although we won't be using most of the names given there. By far the most detailed plant website is the University of California on-line paleobotany lab. The diagrams and hierarchical systems of detail here are most impressive. You may want to scan these pages for the beautiful line drawings alone, but the clear and simple text will trap you. If you want professional detail of fossil plant systematics and distribution, click into the homepage of the International Organisation of Palaeobotany. Berkeley's "Deep Green" website on plant diversity is also very impressive. Finally, here's a ginkgo webpage for everything you want to know about this wonderful tree common on our campus (although it is hardly the only "link between the primitive and modern plants").

Here is our memo describing the Final Examination in this course.

Geology in the News:

I found this very good -- and simple -- YouTube video explaining the basics of evolution. If only more people would watch it. The other videos made by "QualiaSoup" are also worth watching.

With the rupture of an "ice bridge", the Wilkins shelf on the western side of the Antarctic Peninsula appears about to break away, float out into the sea, and melt. This event in itself is negligible on a global scale, but it is more serious evidence that the ice caps are changing rapidly.

Evolution MegaLab is a project just started in Britain to study the recent evolution of the common banded snails found throughout Europe in gardens. The public is being asked to gather samples of snails, along with other data, which will then be used to see how these snails have evolved in the last decade in response to climate patterns and ecological modifications. Cool idea.

Australian and American scientists exploring the Tasman Fracture have discovered several new marine species, including a carnivorous sea squirt (Subphylum Urochordata -- the tunicates, which you remember well) that captures shrimp in a manner like the Venus Fly-Trap. They also recovered 40,000 years worth of coral skeletons which will be valuable for climate research.


April 13-17

Catastrophe: The Cretaceous Mass Extinctions
Recommended Reading: Special readings
Special Events: Osgood Lecture on April 15 (evening).
Assignments: (Preparation questions)

Web Resources for Week #12:

We will spend this week looking at the incredible story of the Cretaceous Mass Extinctions, sometimes called the "K/T" Extinctions (or "KTX" for the cognoscenti). The "Impact Hypothesis" was developed at the University of California, Berkeley, in 1980 when I was there as a graduate student (to my great fortune, even if I had nothing to do with it). It seemed absolutely bonkers at the time, but now we know it as one of those intellectual breakthroughs which changed our perspective on the History of Life. The first website to visit, appropriately, is Berkeley's Cretaceous Extinctions page. You'll love it. It has enough detail to keep us all very busy. Note that dinosaurs were apparently cut down in their prime, although there is still much debate about this. (Read that last article very carefully for a little surprise in it!)

We will emphasize the Impact Hypothesis in this course because I think it is the one best supported by the evidence. (You might like this goofy short YouTube video on the K/T impact story. You'll double the number of viewers!) The volcanic hypothesis is still alive, although in its death struggle. Some of its supporters are bitter, claiming that the K/T extinction argument has been thoroughly corrupted by politics; they even ask for a Congressional inquiry! Seems a bit of sour grapes, but read their page. In contrast with the reasonable (more or less), there are many more exotic ideas about dinosaur extinctions, some tallied here and here. Finally, let's not forget that many more organisms than the dinosaurs were affected by the Cretaceous Extinctions.

If you are considering another geology course, think about Processes & Concepts of Geology (Geo 200) this fall. It is taught by Shelley Judge at 9:30-10:50 a.m. TTh. (Not at 8 a.m.!) It is much fun, and the lab (TTh 1-2:20 p.m.) gives you a chance to see rocks and fossils in more detail. This course is our gateway to the advanced geology courses, but anyone can take it with a 100-level geology course as background. Here is the course webpage for Geology 200.

Here is our memo describing the Final Examination in this course.

Geology in the News:

They take some time to load, but these excellent photographs of the Mt. Redoubt eruption are worth the wait. They come from The Boston Globe via Digg via our own Mícheál O'Duffy.

A very rare -- and very large -- fossil of a giant seabird was found this year in Peru. Note the nasty teeth in the beak, which were apparently bony and very much like those found in Archaeopteryx.

Another important fossil was found recently in Peru: a giant shark in sediments 4-5 million years old. The hypothesis of the associated study is that the famous megalodon sharks and modern white sharks are more distantly related than paleontologists thought.

A bird's egg collected by Charles Darwin himself on the voyage of the HMS Beagle has been rediscovered in the Cambridge University collections. I suppose this is an egg story to coincide with the Easter season. I like that part of Darwin's description of the bird was that its flesh was "most delicately white" when cooked. These properties have dropped out of modern analyses!


April 20-22

Our Cousins at Last: Introduction to Cenozoic Mammals
Recommended Reading: Prothero pages 282-329
Special Events: No class on Friday, April 24, by college order!
Assignments: (Preparation questions)

Web Resources for Week #13:

We had our introduction to mammals back in the Triassic. They were ratty little critters then, for the most part, but now that the dinosaurs are gone our mammals have become very serious. (We now know of a raccoon-sized Mesozoic mammal that ate dinosaurs for lunch, though. albeit small dinosaurs!)

So many mammal groups appeared in the early Cenozoic that we cannot cover them all in this course, or even mention many of them. That diversity dilemma is actually part of the story of the extraordinary adaptive radiation of the mammals, which has been made more complex by recent genetic findings that mammal diversification was not entirely due to the dinosaur extinctions. Note in that story that the three-part division of mammals we see today (monotremes, marsupials and placentals) began in the Cretaceous. This story also introduced me to the "wolf-like cow" Andrewsarchus. Again, the webpages from Berkeley will be highlighted here because they are organized in the most useful ways for us. You will want to look at their short rodents page, and of course make a side visit to the Rat and Mouse Club of America (where one of their ethical principles is that they don't sell their pets to the Snake and Tarantula Club).

Now jump WAY up to the proboscidean (elephants, mammoths, mastodons) page, just to delight in the wondrous variety in modern mammals. There are many pages devoted to mammoths, mastodons and other "Ice Age" mammals, including the Pleistocene exhibits at the Russian Paleontological Institute. Carnivores are always interesting, and are in themselves highly diverse. They include wolves, dogs, cats, raccoons, bears, weasels, hyenas, seals, and walruses. A hero to your cat is the ancient sabretooth, highlighted in special exhibits this year at Berkeley and the Illinois State Museum. You can now make another mammal diversity leap to our unlikely cousins the cetaceans (whales and dolphins; here's an Eocene whale recently found in Egypt). The ungulates, or hoofed mammals, are an excellent group to use in a study of evolutionary patterns. The artiodactyls (cloven-hoofed mammals) include the prosaic sheep, goats, camels, pigs, cows, deer, giraffes, and antelopes (and now whales!). Their relatives the perissodactyls ("odd-hoofed" mammals) include rhinos, horses and tapirs today, but they were much more diverse in the past. The evolution of the horse is a classic story, told well at the "Fossil Horses in Cyberspace" site at the Florida Museum of Natural History. Clearly most of this material will be continued into next week.

Here again is the memo describing the Final Examination on May 4. You will want to read it carefully because it will tell you how to prepare to produce your best test score in this course!

Geology in the News:

There is a new species of ant described from the Amazon which has no males. the female-only populations reproduce entirely by cloning. An interesting evolutionary twist. It is not as unique as the article implies, though. Parthenogenesis, as the process is termed, is not uncommon among animals. Mícheál O'Duffy sent me this story.

You may find this story about the "digital study" of dinosaur skeletons to aid in their reconstruction. The concepts are not at all new, but the computerized tools now used to add flesh to the bones produces amazing results.

Here is a sound-free video (except for the movie trailer at the start!) from the BBC showing a volcanic eruption on one of the Galapagos Islands. Some of the unique flora and fauna on this island is threatened, but such is to be expected when you live on a volcano.


April 27 - May 1

The Wonders of the Thumb: Primate Evolution
Recommended Reading: Allaby pages 68-75; Prothero pages 330-347
Special Events:
Assignments: (Preparation questions)

Web Resources for Week #14:

During this last week we will concentrate on primate and specifically human evolution. We look closely at our own lineage not just because it is of immediate interest to us, but also because we can directly apply concepts of evolutionary theory to organisms intimately familiar to all of us. The Web has almost as many sites on issues of human evolution as it does on dinosaurs, so most of the pages I link you to will provide many other connections. Start first with the Institute of Human Origins, now based at Arizona State University. They have an excellent additional website called "becoming human". For still more detail, try the "Dental Microwear Homepage", which has excellent descriptions and images of the microscopic features paleoanthropologists use to suggest the diets of ancient hominids. Try the interactive "skull module" website where you can take pieces of a human skull and manipulate the images on your screen. Very cool. Think evolution here, especially of the mandibles and cranium. "Evolution's Arrow" is a provocative and stimulating online book by John Stewart (not Jon Stewart!) considering the future of human evolution. You should also pay one last visit to the University of California Museum of Paleontology at Berkeley, which has served us so well on this webpage. Their page on primates is not yet complete, but it does have several links to other resources.

As you examine these websites, please also keep in mind that the fossil record of hominid evolution has produced numerous competing hypotheses, so you will find many disagreements, even on what seem to be basic observations. (Note, for example, the popular pseudoscience surrounding the "aquatic ape hypothesis".) As an example of the hype and expectations which have dogged the study of human evolution for over a century, here is an excellent TalkOrigins webpage on Piltdown Man.

An excellent and clickable hominid-climate evolution diagram is available from the Lamont-Doherty Earth Observatory (Columbia University). Remember that we are not using all the hominid names you find on sites like this.

Here again is the Final Exam Information Sheet. All you need to know about your last test in this course. For review (sort of), you might appreciate the excellent evolution of Homer Simpson!

Geology in the News:

Michael Snader sends us this story of a super-massive "blob" of matter discovered in deepest space which may be one of the youngest objects ever seen. It may represent the beginning of a galaxy when the universe was only 800 million years old. There's the paradox, of course -- this "young" object is now 12.9 billion years old!

Several students sent me this story, with Mícheál O'Duffy first : A remarkable 23-million-year-old fossil was found recently in the Arctic showing a "walking seal" ancestor of the modern seals. Another brilliant example of a transitional fossil. Puijila darwini is its name. It was a fully terrestrial mammal which occasionally may have sought its food in freshwater streams and lakes.

Homo floresiensis (the "Hobbit") lived on the Indonesian island of Flores as early as 17,000 years ago. Controversy rages as to whether this is a newly-discovered hominin species or just another variety of our own Homo sapiens. I have no position yet on this, but I'm moving closer to the conclusion that it is different from us. Now we can look at the skeletons more closely as casts of them have been released for public display.

A black hole has been observed emitting water vapor. Remarkable evidence of just how primordial this stuff is that makes up two-thirds of our bodies.




 May 4 (Monday): Final Examination (7:00 p.m.)


 Notes for the History of Life Course

The History of Life course is a broad overview of the evolution and diversification of life on Earth. It is an introduction to science in general and to evolutionary biology and historical geology in particular, but science majors as well as non-scientists will find the material interesting and challenging. This course, in fact, counts toward the major in geology. It is all great fun, especially when you see that the material not only changes every semester, it changes every week. And wait until you see what strange and marvelous events have happened in the History of Life!

We will meet at 8:00 a.m. Monday, Wednesday, and Friday for lectures in Scovel 105. Attendance is required. You are expected to be in the classroom before 8:00 a.m., especially if there is a quiz. If you have trouble getting out of bed for a morning lecture, this is not the course for you.

The grade in this course will be based on two hourly tests, a final examination, and twelve "pop" (unannounced) quizzes. A student not present in class when a quiz is given (even if it is the beginning of the class) will receive a zero for that assignment. Missed quizzes cannot be "made up" for any reason (except a scheduled and conflicting college event), but the lowest two quiz grades will be dropped at the end of the semester. You will also receive a list of preparation questions before each lecture or accompanying some special reading. Occasionally your written answers to these questions will be collected and graded as a quiz. Please note the dates for all the regularly scheduled tests -- these dates will not be changed except under unusual circumstances. In every class at Wooster you should receive a major graded assignment before the drop date. That major assignment in this course is the first test on February 13. I keep all the grades on a spreadsheet so I can quickly calculate your cumulative course average. It is, however, your responsibility to stay informed about your course performance. Just ask me if you have questions about your scores.

This course has been completed by many students, so there are many notebooks out there filled with pages of lecture details. Because ideas, emphases, and even "facts" change rapidly in biology and geology, use old class notes with caution.

It is possible to take this course "S/NC", unless you plan to major in geology. The minimum letter grade for an "S" in this course is C, which is a final average better than 75% on tests and quizzes. I do not encourage the S/NC option -- usually students taking it earn high grades anyway, or they fail with scores below C.

This course is administered under the Code of Academic Integrity, which you should review in your copy of The Scot's Key. Violations of the Code in this course include consulting notes, other student papers, or electronic devices during tests and quizzes. My penalties for cheating start with an F in the course and then get worse.

The faculty of the College has recently approved a new policy regarding conflicts between extracurricular and academic events. I agree with it and will simply provide the suggested syllabus statement from the discussion: "The College of Wooster is an academic institution and its fundamental purpose is to stimulate its students to reach the highest standard of intellectual achievement. As an academic institution with this purpose, the College expects students to give the highest priority to their academic responsibilities. When conflicts arise between academic commitments and complementary programs (including athletic, cultural, educational, and volunteer activities), students, faculty, staff, and administrators all share the responsibility of minimizing and resolving them. As a student you have the responsibility to inform the faculty member of potential conflicts as soon as you are aware of them, and to discuss and work with the faculty member to identify alternative ways to fulfill your academic commitments without sacrificing the academic integrity and rigor of the course."

 

Textbooks

Our "textbooks" are different this semester. My students and I have grown tired of our expensive and not very effective traditional textbooks, so this semester we are going to use a colorful encyclopedia and a popular book on evolution. I figure I'm saving you at least $110, and you are guaranteed to find these books especially interesting. This webpage and our class lectures will give us the narrative thread for the course, so pay close attention! We're also going to test the hypothesis that this encyclopedia can become a common resource for all the introductory geology courses. The readings are scattered throughout the books.

Allaby, M. et al., 2008. The Encyclopedia of Earth: A Complete Visual Guide. University of California Press (Berkeley); 608 pages; ISBN 978-0520254718. We will not directly use about half of this book, but I bet you can't help browsing through all of it! [Course readings are listed in the lecture outline as "Allaby".]

Prothero, D.R., 2007. Evolution: What the Fossils Say and Why It Matters. Columbia University Press (New York); 408 pages; ISBN 978-0231139625. This book is written as a strong and effective defense of evolution against the modern forms of creationism. We will not be talking directly about creationism, but I hope you read the sections which cover it very closely. [Course readings are listed in the lecture outline as "Prothero".]

Grading Format

Testing Date Percentage of Grade Notes
1st Exam February 13  20%
2nd Exam April 3  25%
Final Exam May 4, 7:00 p.m.  30%
Quizzes (Pop!) (Unannounced)  25% total Lowest two dropped

 

Teaching Assistant

Jesse Davenport, a junior geology major, is our teaching assistant this semester. Jesse will also serve as a tutor for this course.

 

Course Instructor

I have an appointment schedule posted outside my office door in Scovel. Please sign up for an appointment if you have any questions or just want to talk. In a large course such as this students often think I am too busy to see them individually unless there is a problem. This is not true. Please visit me if there is anything at all you wish to discuss. The material for this course is too interesting to let it go by as just a series of lectures!

Mark A. Wilson
Scovel 120
Telephone ext. 2247
mwilson@wooster.edu

Pteranodon


Pteranodon by Heinrich Harder (1916)


Science, Evolution, and Creationism ...
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