The Keck Ohio Project

Late Ordovician Paleontology, Sedimentology
and Stratigraphy in Ohio, Indiana and Kentucky


A Keck Geology Consortium Project
June 19 - July 16, 1999




Bioimmuration and what it tells us about the systematics and paleoecology of encrusting organisms in the Cincinnatian Group (Upper Ordovician, Ohio, Indiana, Kentucky)


Woody Fischer
Department of Geology, The Colorado College
Colorado Springs, CO 80903

In certain environments there are organisms which undergo rapid postmortem decay or dissolution and are rarely (perhaps never) represented in fossil assemblages. Lagerstatten are examples of where unique fossil preservation has occurs. One commonly overlooked process of preservation is bioimmuration, whereby soft-bodied organisms and those with aragonite shells are immured in the calcitic shells of others (Taylor 1990).

Bioimmurations are an incredible resource for research; however this study is not simply limited to just describing immured organisms. For example, if Ordovician bryozoans encrusted an aragonitic bivalve, then whether the bryozoans formed a cast or mold of the immured organism, the underside of the bryozoans would be exposed after bivalve dissolution. With their undersides now revealed, we can develop insight into ecological relationships (such as overgrowth and competition) not often seen in paleoecology.

My hypothesis is that repeatable ecological relationships exist between types of hosts and encrusters, which reveal competition between encrusters, and which depict a predictable succession between encrusters. My hypothesis will also extend into looking into the connection between types of hosts and succession of encrusters in situations where there has been soft-bodied preservation. Results derived from this working hypothesis will be directly connected from paleontological relationships to ecological theory, offering a valuable look not simply at bioimmuration phenomena, but also at the ecology of encrusting processes.




Woody Fischer


An unusual boring or bioimmuration incised in the bottom of a bioimmuring bryozoan from the Whitewater Formation, one mile south of Richmond, Indiana. The substrate for the original bryozoan was the bivalve Ambonychia.


A bryozoan bioimmured this valve of the clam Ambonychia, and then it appeared to overgrow its own growing surface after dissolution of the clam's aragonitic shell. This could be important evidence for the timing of aragonite dissolution in this Ordovician "calcite sea".


Woody Fischer, always a well-dressed field geologist, examining the evidence.

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