A phylogenetic study of the nymphalid butterflies and their host plant affinities in the Florissant Formation (Late Eocene, Colorado)
Amanda Cook
Department of Geology, The College of Wooster
Wooster, Ohio 44691

The Florissant Fossil Beds were established as a national monument by the National Park Service in 1969 to preserve the Late Eocene plant and insect fossils found there today. The fine-grained, laminated lake shales formed by eruptions from the Thirtynine Mile volcanic field contain exceptionally well preserved fossils. Florissant 34 m.y. ago was very different than it is today. This lake environment was host to a very diverse plant and insect community. Warm temperate to sub tropical temperatures and an elevation close to what it is today (8500 ft) has been calculated using the plant fossils. Florissant has produced the most diverse fossil butterfly collection known to date, with 12 species. Given this rare opportunity to look at both fossil butterflies and host plants, I plan to compose a list of host plants for the butterflies in the Florissant Formation belonging to the Family Nymphalidae, which is the best represented of the families. Because so much study has been done in the classification of insects, we are able to assign a recent counterpart to each fossil genus. This year I hope to draw some phylogenetic relationships between fossil and recent genera of the nymphalids so that I can get a clearer picture as to what significant evolutionary changes occurred. With this I hope to avoid the problem of assuming that recent nymphalids live the same way in their environment as they did 34 million years ago.  

 

Amanda Cook is a senior geology major at The College of Wooster. Her hometown is Cleveland, Ohio. Her research this summer was made possible by an internship at the Florissant Fossil Beds National Monument. Her field supervisor was Dr. Herb Meyer, a paleobotanist with the National Park Service.

Prodryas persephone Scudder, 1878. Harvard Museum of Comparative Zoology; photo- graph by Herb Meyer. Fine divisions of the scale are millimeters.

 

Chlorippe wilmattae Cockerell, 1907. Harvard Museum of Comparative Zoology; photo- graph by Herb Meyer. Fine divisions of the scale are millimeters.

 

Porana tenuis (unpublished). Family Convolvulaceae, host plant of Juptellia sp.; University of Florida Museum; photograph by Herb Meyer. Fine divisions of the scale are millimeters.

 

Humulus sp. (unpublished). Family Cannabidaceae, host plant of Vanessa sp.; University of California Museum of Paleontology; photograph by Herb Meyer. Fine divisions of the scale are millimeters.

 

Link to homepage of the Florissant Fossil Beds National Monument.

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